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You need a computer that can run at least Java 1.7 to use the full Jalview application (unfortunately this excludes users with older Macs - sorry). If java is installed, then the 'Launch Jalview Desktop' link at the top should work (if your browser asks you to save the application-x-jnlp file or choose an application to run it with, look for 'Java Web Start', or locate your java installation directory and look for the javaws binary).